Thursday, January 22, 2009

Population Ecology II

This section on population ecology is the topic that I am most concerned about teaching via distance. It seems easier to show you in person how I work problems than it is to try to explain in writing how to work the problems. In addition, I have always found it easier to teach math and graphing when I can give students some problems to solve and I can walk around the classroom peering over their shoulders while they work (it is fun to freak out freshmen that way!).

After you are comfortable with the paremeters that I introduced in teh Population Ecology I. blog, then I would read the articles from the EoE in the following order.

Population ecology
Exponential growth
Logistic growth
Carrying Capacity
Intraspecific competition

These articles (which I have written) attemtp to introduce the readers to the two most important mathematical models that have been used to describe simple population growth.

Exponential Growth

From the first lesson on Population Ecology we learned that the population growth rate (dN/dt) can be calculated as the product of the per capita growth rate (r) and the population size (N).

dN/dt = rN

This is the fundamental equation describing population growth and this equation is always true.

If we want to use this equation to analyze how population sizes change over time, then it makes sense to start by examining the simplest formulation of this equation which occurs when the per capita growth rate is constant. The equation dN/dt = rN when r is constant is known as the exponential growth equation and this equation describes a patter on growth known as exponential growth.

The graph plotting how population size changes over time is shown in the Exponential Growth article. This graph shows an exponential growth curve (sometimes known as the "j-curve"). If you have questions about why the graph has this shape let me know and I will try to explain it more thoroughly.

It is important that you are able to look at this graph and determine all of the information held in the graph. The exponential growth curve allows us to discuss how two parameters change over time- 1) the population size (shown by the x-axis) and 2) the population growth rate (shown by the slope of the line). I find that it is easier to discuss only one parameter at a time so let's start with the population size.

1) Over time, the population size increases (we know this because the line has a positive slope).

Now let's think about the population growth rate.

2) Over time, the population growth rate increases (we know this becasue the line gets steeper over time.

3) Over time, the rate at which the population growth rate increases over time, increases over time (we know this because the slope increases faster and faster over time).

Thus, if populations are growing exponentially then they keep increasing in size at an ever faster rate forever and ever.

Now try this-

Can you draw the following graphs?

1) plot how the population growth rate varies over time.

(hint- we have alredy described what this pattern will look like using words- just turn these words into pictures).

2) plot how the population growth rate depends on population size.

(hint- this graph is a little trickier, but we do have an equation that relates the two variables)

3) plot how the per capita growth rate varies over time.

(hint- think about what the basic assumption we made aboiut exponential growth)

4) plot how the per capita growth rate varies over time.

(see the hint from number 3)


Exponential Growth is Unrealistic

Because population sizes keep increasing at ever faster rates for ever, exponential growth does not seem to be an accurate description of population growth in most animals, plants, and microbes. If this is an unrealistic model then why did I teach it to you? I started with exponential growth becasue it is the simplest model of population growth and scientists always like to describ the world using the simplest models that they can.

Obviously, in this case we have started with a model that is too simple to realistically describe the world. What is wrong with the exponential growth model? The fundamental assumption we made about exponential growth is that the per capita growth rate is constant. This must not be a realistic assumtpion.

It is important that you understand, and are able to explain, both the mathematical reasons and biological reasons that exponential growth is an unreasonable model of population growth. I tried to explain biologically why exponential growth is unrealistic in the "Exponential Growth" article and the attached Powerpoint presentation so take a look at those.

Powerpoint presentation "Why is Exponential Growth Unrealistic?" http://www.slideshare.net/secret/IDPugQtl2wvONv


Final Thoughts

Many students find using math to think about biological concepts and using graphs to illustrate pattersn to be difficult. It is probably difficult for students because they have not had very much practice doing it. If you are comfortable using math and graphs, then most of what we are doing will not be too difficult. However, if you lack a lot of experience using math and graphs, this section might be a bit frustrating. My advice to you is to keep plugging away. Once you learn how to approach these problems, then you will find that you have developed a skill that you can use over an over again. It will, however, require some practice to develop these skills. Please let me know if you are having any problems or questions. You can post on the blog, send me an email, or if you think it will help to actually talk, then we can talk over the phone (it is frustratingly difficult to quickly and easily show you graphs electronically). I need to go educate the masses of Texas Tech, but I will be back on-line soon to talk about logistic growth (a more realistic and useful model of population growth).

4 comments:

  1. Here are my thoughts...

    first my computer was not loving the link to the slideshow which is odd because it has let me watch the other on desert adaptations. I will give it another try later.

    As to the graphs...

    1) This would be a "j" curve where the x-axis is time (t) and the y-axis is population (N)

    2) This graph would have as the x-axis population size (N) and the y-axis as the population growth rate. There would be two slopping lines (one positive and one negative) and their intersection would be the carrying capacity

    3) Here I begin to falter...this is a pure guess...would this be the "S" curve. I know that the per capita growth rate is different than the population growth rate, but I am a bit foggy on this!

    4) I don't understand what the two variables are here. One would be the per capita growth rate, but what is the other?

    Any words of wisdom would be appreciated. Hope all is going well for everyone!

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  2. Brandy,

    I am sorry that you are having trouble with the slideshow. They have all worked for me. Sometimes I have had trouble with the slide share website, but eventually they get things fixed. Let me know if you continue to have trouble seeing the slideshows.

    1) Your first answer is correct.

    2) The graph asked for in question 2 is shown as Exponential Growth Graph #2 in the Fun With Graphs Exponential Growth slideshow.

    Because dN/dt = rN, dN/dt changes proportionally with N. The graph asked for in question 2 is another j-curve where the y axis population growth rate and the x axis is time.

    3 & 4) You are 100% correct that question 4 made no sense because I originally didn't finish the statement. Questio 4 now reads-plot how the per capita growth rate varies over time (I left out the over time part origninally, sorry)

    Please take a look at the Population Ecology Basic Parameters to review the relationship between the "population growth rate" and the "per capita growth rate". "Per capita" rates are averaged across all members of the population so to calculate a per capita rate you always divide the population rate by population size.

    any per capita rate = the population rate/N

    Thus, the population growth rate measures the change in population size over time (units- individuals/time) and the per capita growth rate measures the change in population over time averaged across all individuals (units-individuals/time/individual)

    The graphs I asked you to draw in questions 3 and 4 don't make much sense if you are still a little unsure about what the per capita growth rate is. The difference between the population growth rate and the per capita growth rate is a potentially confusing (but important) issue. Take a look at the Population Ecology, Basic Parameters section and let me know if you have more questions.

    Here is a hint about how to draw the two graphs- what is the critical assumption made in exponential growth?

    I hope this helps. Please keep sending me questions and comments,

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  3. OK I was able to get into the slideshow. This makes more sense now. THANKS!

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  4. This is annoying! Sorry the computer logged me in as Karlton again...ARGH!

    ReplyDelete